diff --git a/core/chromosome.cpp b/core/chromosome.cpp
index 5652604c..8e93562d 100644
--- a/core/chromosome.cpp
+++ b/core/chromosome.cpp
@@ -2348,60 +2348,6 @@ void Chromosome::PrintMutationRunExperimentSummary(void)
 	}
 }
 
-#if (SLIMPROFILING == 1)
-// PROFILING
-#if SLIM_USE_NONNEUTRAL_CACHES
-void Chromosome::CollectMutationProfileInfo(void)
-{
-	Chromosome &chromosome = TheChromosome();
-	
-	// maintain our history of the number of mutruns per haplosome and the nonneutral regime
-	profile_mutcount_history_.emplace_back(mutrun_count_);
-	profile_nonneutral_regime_history_.emplace_back(last_nonneutral_regime_);
-	
-	// track the maximum number of mutations in existence at one time
-	int registry_size;
-	
-	population_.MutationRegistry(&registry_size);
-	profile_max_mutation_index_ = std::max(profile_max_mutation_index_, (int64_t)registry_size);
-	
-	// tally up the number of mutation runs, mutation usage metrics, etc.
-	int64_t operation_id = MutationRun::GetNextOperationID();
-	
-	for (std::pair<const slim_objectid_t,Subpopulation*> &subpop_pair : population_.subpops_)
-	{
-		Subpopulation *subpop = subpop_pair.second;
-		std::vector<Haplosome *> &subpop_haplosome = subpop->parent_haplosomes_;
-		
-		for (Haplosome *haplosome : subpop_haplosome)
-		{
-			const MutationRun **mutruns = haplosome->mutruns_;
-			int32_t mutrun_count = haplosome->mutrun_count_;
-			
-			profile_mutrun_total_usage_ += mutrun_count;
-			
-			for (int32_t mutrun_index = 0; mutrun_index < mutrun_count; ++mutrun_index)
-			{
-				const MutationRun *mutrun = mutruns[mutrun_index];
-				
-				if (mutrun)
-				{
-					if (mutrun->operation_id_ != operation_id)
-					{
-						mutrun->operation_id_ = operation_id;
-						profile_unique_mutrun_total_++;
-					}
-					
-					// tally the total and nonneutral mutations
-					mutrun->tally_nonneutral_mutations(&profile_mutation_total_usage_, &profile_nonneutral_mutation_total_, &profile_mutrun_nonneutral_recache_total_);
-				}
-			}
-		}
-	}
-}
-#endif
-#endif
-
 
 //
 // Eidos support
diff --git a/core/species.cpp b/core/species.cpp
index 906f3c0d..1e43cc70 100644
--- a/core/species.cpp
+++ b/core/species.cpp
@@ -3031,6 +3031,60 @@ void Species::ReturnShuffleBuffer(void)
 	shuffle_buf_borrowed_ = false;
 }
 
+#if (SLIMPROFILING == 1)
+// PROFILING
+#if SLIM_USE_NONNEUTRAL_CACHES
+void Species::CollectMutationProfileInfo(void)
+{
+	Chromosome &chromosome = TheChromosome();	// only keeping the history for the first chromosome right now, should keep it for all
+	
+	// maintain our history of the number of mutruns per haplosome and the nonneutral regime
+	profile_mutcount_history_.emplace_back(chromosome.mutrun_count_);
+	profile_nonneutral_regime_history_.emplace_back(last_nonneutral_regime_);
+	
+	// track the maximum number of mutations in existence at one time
+	int registry_size;
+	
+	population_.MutationRegistry(&registry_size);
+	profile_max_mutation_index_ = std::max(profile_max_mutation_index_, (int64_t)registry_size);
+	
+	// tally up the number of mutation runs, mutation usage metrics, etc.
+	int64_t operation_id = MutationRun::GetNextOperationID();
+	
+	for (std::pair<const slim_objectid_t,Subpopulation*> &subpop_pair : population_.subpops_)
+	{
+		Subpopulation *subpop = subpop_pair.second;
+		std::vector<Haplosome *> &subpop_haplosome = subpop->parent_haplosomes_;
+		
+		for (Haplosome *haplosome : subpop_haplosome)
+		{
+			const MutationRun **mutruns = haplosome->mutruns_;
+			int32_t mutrun_count = haplosome->mutrun_count_;
+			
+			profile_mutrun_total_usage_ += mutrun_count;
+			
+			for (int32_t mutrun_index = 0; mutrun_index < mutrun_count; ++mutrun_index)
+			{
+				const MutationRun *mutrun = mutruns[mutrun_index];
+				
+				if (mutrun)
+				{
+					if (mutrun->operation_id_ != operation_id)
+					{
+						mutrun->operation_id_ = operation_id;
+						profile_unique_mutrun_total_++;
+					}
+					
+					// tally the total and nonneutral mutations
+					mutrun->tally_nonneutral_mutations(&profile_mutation_total_usage_, &profile_nonneutral_mutation_total_, &profile_mutrun_nonneutral_recache_total_);
+				}
+			}
+		}
+	}
+}
+#endif
+#endif
+
 
 //
 // TREE SEQUENCE RECORDING
diff --git a/eidos/eidos_globals.h b/eidos/eidos_globals.h
index caf25a8f..93f3d5fe 100644
--- a/eidos/eidos_globals.h
+++ b/eidos/eidos_globals.h
@@ -34,6 +34,10 @@
 #include <unordered_map>
 #include <cstdint>
 
+// Workaround for Xcode bug: when you want to debug build problems with a Release build related to profiling, uncomment this,
+// since the target-level definition of SLIMPROFILING doesn't seem to affect syntax highlighting and build errors correctly.
+//#define SLIMPROFILING 1
+
 #if defined(__APPLE__) && defined(__MACH__)
 // On macOS we use mach_absolute_time() for profiling and benchmarking
 #include <mach/mach_time.h>