diff --git a/src/Sortable.js b/src/Sortable.js
index 47761808e..d2fba8e41 100644
--- a/src/Sortable.js
+++ b/src/Sortable.js
@@ -336,7 +336,7 @@ let nearestEmptyInsertDetectEvent = function(evt) {
 
 let _checkOutsideTargetEl = function(evt) {
 	if (dragEl) {
-		dragEl.parentNode[expando]._isOutsideThisEl(evt.target);
+		dragEl.parentNode && dragEl.parentNode[expando]._isOutsideThisEl(evt.target);
 	}
 };
 
@@ -774,7 +774,7 @@ Sortable.prototype = /** @lends Sortable.prototype */ {
 				parent = target;
 			}
 
-			dragEl.parentNode[expando]._isOutsideThisEl(target);
+			dragEl.parentNode && dragEl.parentNode[expando]._isOutsideThisEl(target);
 
 			if (parent) {
 				do {
@@ -1101,7 +1101,7 @@ Sortable.prototype = /** @lends Sortable.prototype */ {
 
 			// no bubbling and not fallback
 			if (!options.dragoverBubble && !evt.rootEl && target !== document) {
-				dragEl.parentNode[expando]._isOutsideThisEl(evt.target);
+				dragEl.parentNode && dragEl.parentNode[expando]._isOutsideThisEl(evt.target);
 
 				// Do not detect for empty insert if already inserted
 				!insertion && nearestEmptyInsertDetectEvent(evt);
@@ -1327,6 +1327,13 @@ Sortable.prototype = /** @lends Sortable.prototype */ {
 			if (el.contains(dragEl)) {
 				return completed(false);
 			}
+		} else {
+			// unmounted dragEl from previous target who is dragging over a new target but can not be dropped
+			if (putSortable && putSortable.lastPutMode !== 'clone') {
+				rootEl.appendChild(dragEl);
+			} else if (rootEl !== dragEl.parentNode) {
+				dragEl.parentNode && dragEl.parentNode.removeChild(dragEl);
+			}
 		}
 
 		return false;